Publicaciones 2008-2017


  • (39) Pieterse W, Benítez HA, Addison P. 2017. The use of geometric morphometric analysis to illustrate the shape change induced by different fruit hosts on the wing shape of Bactrocera dorsalis and Ceratitis capitata (Diptera: Tephritidae). Zoologischer Anzeiger - A Journal of Comparative Zoology 269: 110-116.(PDF)

  • (38) Lunardi R, Benítez HA, Câmaraa TP, Gomes LP, Arrais-Silva W. 2017. Head shape variation in response to diet in Triatoma williami (Hemiptera, Reduviidae: Triatominae), a possible Chagas disease vector of legal Amazônia. Zoologischer Anzeiger - A Journal of Comparative Zoology. 267(3): 187-193.(PDF)

  • (37) Benítez HA, & Vargas HA. 2017. Sexual dimorphism and population differentiation in the Chilean Neotropical moth Macaria mirthae (Lepidoptera, Geometridae): a wing geometric morphometric example. Revista Brasileira de Entomologia 61(4): 365-369




  • (36) Mikac KM, Lemic D, Bažok R, Benítez HA. 2016. Wing shape changes: a morphological view of the Diabrotica virgifera virgifera European invasion. Biological Invasion 18(12): 3401–3407. DOI: 10.1007/s10530-016-1252-9.

  • (35) Lemic D, Benítez HA, Püschel T, Virić Gašparić H, Šatvar M, Bažok R. 2016. Ecological morphology of the sugar beet weevil Croatian populations: evaluating the role of environmental conditions on body shape. Zoologischer Anzeiger 260(1): 25–32. 

  • (34) Lemic D, Mikac KM, Kozina A, Benítez HA, Bazok R. 2015. Monitoring techniques of the western corn rootworm are the precursor to effective IPM strategies. Pest Management Science 72(2) 405-417. DOI: 10.1002/ps.4072.



  • (33a) Benítez HA., Vargas H, & Püschel TA. 2015 Left-Right Asymmetry and morphological consequences of a host shift in the oligophagous Neotropical moth Macaria mirthae (Lepidoptera: Geometridae). Journal of Insect Conservation 19(3): 589-598.
    (33b) Erratum to: Left–right asymmetry and morphological consequences of a host shift in the oligophagous Neotropical moth Macaria mirthae (Lepidoptera: Geometridae)  19(3): 599. 

  • (32) Zúñiga-Reinoso A, Benitez HA. 2015. The overrated use of the morphological cryptic species concept: An example with Nyctelia darkbeetles (Coleoptera: Tenebrionidae) using geometric morphometrics. Zoologischer Anzeiger - a Journal of Comparative Zoology 255:47-53.

  • (31) Schmieder  DA., Benítez HA., Borissov IM & Fruciano C. 2015. Bat Species Comparisons Based on External Morphology: A Test of Traditional versus Geometric Morphometric Approaches. PLOS ONE, 10.1371/journal.pone.0127043.



  • (30) Püschel T, Espejo J, Sanzana MJ  & Benítez HA. 2014. Analysing the floral elements of the lost tree of Easter Island: A morphometric comparison between the remaining ex-situ lines of the endemic extinct species Sophora toromiro. PLOS ONE; DOI: 10.1371/journal.pone.0115548.

  • (29) Benítez HA, Püschel T, Lemic D, Čačija M, Kozina A, Bažok R. 2014. Ecomorphological variation of the wireworm cephalic capsule: Studying the interaction of environment and geometric shape. PLOS ONE. Jul 8;9(7):e102059. doi: 10.1371/journal.pone.0102059.

  • (28) Benítez HA, Lemic D, Püschel T, Bravi R, Buketa M, Bažok R. 2014. Morphological Integration and Modularity in Diabrotica virgifera virgifera LeConte (Coleoptera: Chrysomelidae) hind wings. Zoologischer Anzeiger - A journal of comparative Zoology 253(6): 461-468.

  • (27) Benitez HA, Lemic D, Bažok R, Gallardo-Araya C, Mikac KM (2014) Evolutionary Directional Asymmetry and Shape Variation in Diabrotica v. virgifera (Coleoptera: Chrysomelidae): an example using hind wings. Biological Journal of Linnean Society 111(1): 110-118.

  • (26) Püschel T. & Benítez HA. 2014. Femoral Functional Adaptation: a comparison between Hunter-Gatherers and Agriculturalists using Geometric Morphometrics. International Journal of Morphology 32(2): 627-63.

  • (25) Storey-Palma, J., Benítez, HA, Mundaca, EA & Vargas HA. 2014. Egg laying site selection by a host plant specialist leaf miner moth at two intra-plant levels in the northern Chilean Atacama Desert. Revista Brasileira de Entomología. 58(3): 280-284.

  • (24) Benitez HA. 2014. Is the Fever for High Impact a Disadvantage for Systematists. Neotropical Entomology DOI 10.1007/s13744-014-0217-x  Editorial Letter 

  • (23) Lemic D, Benítez HA, Bažok R. 2014. Intercontinental effect on sexual shape dimorphism and allometric relationshisps in the beetle pest Diabrotica virgifera virgifera LeConte (Coleoptera: Chrysomelidae). Zoologischer Anzeiger 253(3): 203–206.

  • (22) Benítez HA, Püschel T. 2014. Modelando la varianza de la forma: Morfometría geométrica aplicaciones en Biología Evolutiva. International Journal of Morphology. 32(3):998-1008. 

  • (21) Benítez HA, Pizarro-Araya J, Bravi R, Sanzana MJ, Alfaro F (2014). Morphological variation on isolated populations of Praocis (Praocis) spinolai Gay & Solier, 1840 (Coleoptera: Tenebrionidae) Journal of Insect Science 14(11): 1-12.

  • (20) Vergara O, Benítez HA, Pincheira M, Jerez V. 2014. Determinación del dimorfismo sexual en la forma corporal de Chiasognathus grantii (Coleoptera: Lucanidae). Revista Colombiana de Entomologia. 40(1): 104-10.



  • (19) Sanzana MJ, Parra LE, Sepulveda E, Benítez HA. 2013. Latitudinal gradient effect on the wing geometry in Auca coctei Guérin (Lepidoptera, Nymphalidae). Revista Brasileira de Entomologia. 57(4): 411-416; doi: 10.1590/S0085-56262013005000045

  • (18) Benitez HA, Bravi R, Parra LE, Sanzana MJ, Sepúlveda-Zúñiga E. 2013. Allometric and non-allometric pattern in sexual dimorphism discrimination of wing shape in the parasitoid, Ophion intricatum: do two male morphotypes coexist? Journal of Insect Science 13: 143. 1-10

  • (17) Briones RC, Jerez V, Benítez HA. 2013. Vertical Diversity of Beetles (Insecta: Coleoptera) Associated with Lithraea caustica (Anacardiaceae) in Patches of Sclerophyllous Forest in Central Chile. Journal of the Entomological Research Society. 15(3): 41-52.

  • (16) Benitez HA, Avaria-Llautureo J, Canales-Aguirre CB, Jerez V, Parra L, Hernandez CE. 2013 Evolution of sexual size dimorphism and its relationship with sex ratio in carabid beetles of Genus Ceroglossus Solier. Current Zoology 59(6): 769-777.

  • (15) Vargas HA, Benítez HA. 2013. Egg phenology of a host-specialist butterfly in the western slopes of the northern Chilean Andes. Neotropical Entomology DOI 10.1007/s13744-013-0170-0.

  • (16) Bravi R & Benítez HA. 2013. Left-Right asymmetries and shape analysis on Ceroglossus chilensis (Coleoptera: Carabidae). Acta Oecologica 52: 57-62.

  • (13) Benitez HA., Sanzana MJ., Jerez V., Parra L., Hernández CE. & Canales-Aguirre CB. 2013. Sexual shape and size dimorphism in carabid beetles of the Genus Ceroglossus. Is the geometric body size similar between sexes due to sex ratio? Zoological Science 30(4): 289-295 doi: 10.2108/zsj.30.289-

  • (12) Benitez HA. 2013. Assessment of patterns of fluctuating asymmetry and sexual dimorphism in carabid body shape. Neotropical Entomology 42(2): 164-169.

  • (11) Benitez HA. 2013, Sexual dimorphism using geometric morphometric approach. In Moriyama H. Sexual Dimorphism ISBN 980-953-307-175-1 Intech. Rijeka, Croatia.

  • (10) Alarcón ME., Huang C-G., Dubey AK. & Benitez HA. 2013. A gregarine from the gut of cat flea, Ctenocephalides felis (Bouché) (Siphonaptera: Pulicidae) in Taiwan: dynamic of infection patterns. Veterinary Parasitology 192 (1-3) 51-56. 




  • (9) Benítez HA & Parra LE. 2012. Asimetría Fluctuante: Una herramienta morfo-funcional para medir estabilidad del desarrollo. International Journal of Morphology. 29(4).

  • (8) Sanzana MJ, Parra LE, Benítez HA & Espejo J. 2012. Entomofauna Polinizadora de Eucalyptus nitens en los Huertos Semilleros de la Hacienda Rucamanqui, Huépil, Región del Biobío, Chile Central. Bosque 33(1).

  • (7) Sepulveda E, Parra LE, Benítez H, & Rojas C. 2012 Estados de naturalidad y heterogeneidad vegetacional de humedales palustres y su efecto sobre la diversidad de macrolepidópteros. SHILAP Revista de lepidopterología 40:158 - 155-170.

  • (6) Storey-Palma J., Benitez HA., Parra LE., Vargas HA.  2012. Identification of sap-feeder instars in Angelabella tecomae Vargas & Parra (Lepidoptera, Gracillariidae) on Tecoma fulva fulva (Bignoniaceae).  Revista Brasileira de Entomologia 56(4): 508-510



  • (5) Benítez H. Briones R. & Jerez V 2011. Intra and Inter-population morphological variation of shape and size of Ceroglossus chilensis (Eschscholtz, 1829) in Chilean Patagonia.  Journal of Insect Science 11:98 1-9.

  • (4) Benítez H., Parra L. E, Sepulveda E & Sanzana MJ. 2011. Geometric perspectives of sexual dimorphism in the wing shape of Lepidoptera: the case of Synneuria sp. (Lepidoptera: Geometridae). Journal of the Entomological Research Society 13(1):53-60.



  • (3) Benítez H., M. Vidal, R. Briones & V. Jerez. 2010 Sexual Dimorphism and Morphological Variation in Populations of Ceroglossus chilensis (Eschscholtz, 1829) (Coleoptera: Carabidae) . Journal of the Entomological Research Society 12(2) 87-95.

  • (2) Benítez H, Jerez V & Briones R. 2010. Proporción sexual  y Morfometría  para dos poblaciones de Ceroglossus chilensis (eschscholtz, 1829) (Coleoptera: Carabidae) en la Región del Bio bío Chile. Revista Chilena de Entomología. 35 (1): 61-70.



  • (1) Benítez. H., R. Briones & V. Jerez. 2008. Asimetría Fluctuante en dos poblaciones de Ceroglossus chilensis (Coleoptera Carabidae) en el agroecosistema Pinus radiata, Región del BioBío. Gayana72(2): 131-139.